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do they do that to get the most power or does it matter with the gloves they wear?
You punch with those knuckles because they are larger and the musculature and support structures are larger on the thumb side of the hand. This makes it less likely that you will break your hand when hitting with the first 2 knuckles; the extra power is just incidental.
No fighter would be dumb enough to intentionally hit with the last 2 knuckles.
It depends on the fighter. Chuck Liddel, for instance, punches almost exclusively with his thumb.
It depends where you're coming from in terms of training.
If you read Jack Dempseys book he goes with the last 3 knuckles . He suggest a test where you line them up vertically against a wall and it shows the direct line from your shoulder to your knuckles. Thus not breaking the line of power and improtantly not breaking your wrist.
A lot of Karate based styles foucs on the first 2 knuckles.
Also if you're striking with a vertical fist or straight fist you will notice the difference in the way punching influences which knuckles will strike
I'm on Dempsey's side for this one. I find that aiming with the 1st two knuckles is mechanically less efficient than aiming with the middle two. The 1st two might be bigger, but I find I need to tilt my hand in a way that makes the alignment less efficient..
I'd say that you should strive to hit with a variety of different points of impact, varying punch by punch. For straight punches i'd say that my first 2 knuckles generally make impact, whereas uppercuts and hooks generally land on the last 3.
The danger with only landing with your first 2 knuckles is that it can lead to boxer's knuckle. Conversely, trying to land with the other 3 all the time can throw your hand out of alignment and potentially lead to breaking your wrist. With gloves & hand wraps on, it becomes less of a problem. But, in general i'd say use whatever feels comfortable for you. I try to use a combination of the 2
I feel it's actually more likely with first 2 knuckles making contact. Your wrist is much more stable with last 3 making contact. Try doing pushups with 2 vs 3 knuckles and not how much more stable your wrist is.
I feel more stability when punching a bag bare-knuckle with flat surface of last 3 knuckles. I've had minor sprains in my pinkie couple of times, but broke my index when I did overhand into the elbow, in sparring (with 16 oz glove, lol). I've had 7 am MMA fights if that's relevant.