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Don't think centre backs, you can get by there with very poor athleticism if you have good positional play...and it doesn't require a whole lot of running. Modern day wing backs though, for sure are some of the best athletes in soccer.
There are some centre backs that can flat out fly and be agile. They are pretty tall be normal human standards. In modern day top flight soccer, the traditional centre forward or target man is kind of disappearing. There is a really good article by ESPN about Giroud and Mandzukic as they meet in the WC final. The centre backs would usually defend against tall strikers who are like equivalent to BBall centers and PFs, but thats changing
http://www.espn.com/soccer/fifa-wor...atia-owe-a-lot-to-old-school-giroud-mandzukic
The centre backs nowadays have to do a lot of chasing around and be very agile. These guys are also the tallest players on the team too often than not.